SecReT4
T4SS ID262
StrainCupriavidus metallidurans CH34
Repliconchromosome [Browse all T4SS(s) in this replicon]
AccessionNC_007973
Location1378405..1378992; 1448292..1460364
NameTrb
Functionconjugation
ClassificationType IVA; Type P
sequence This T4SS has been mentioned in a genome sequencing report.

T4SS components
ComponentTrbBTrbCTrbDTrbETrbFTrbGTrbHTrbITrbJ
Number111111011
ComponentTrbKTrbLTrbNTraFTraG
Number01011

The information of T4SS components from NC_007973
Region 1: 1378405..1378992
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#Locus tag (Gene)Coordinates [+/-], size (bp)Protein GIProductComponent
1Rmet_1251 (tnp)1373579..1374580 [-], 100294310195transposase ISRme5 (copy a, CMGI-2) 
2Rmet_12521374596..1375468 [+], 87394310196hypothetical protein 
3Rmet_12531375516..1375833 [+], 31894310197putative dna binding protein 
4Rmet_1254 (repA)1375860..1376717 [+], 85894310198replication initiator and transcription repressor protein 
5Rmet_1255 (parA)1376971..1377609 [+], 63994310199ATPase involved in plasmid partitioning 
6Rmet_1256 (parG)1377606..1377878 [+], 27394310200regulator of plasmid copy number 
7Rmet_12571377875..1378408 [+], 53494310201hypothetical protein 
8Rmet_1258 (traF)1378405..1378992 [+], 58894310202plasmid conjugal transfer transmembrane protein  TraF
9Rmet_1259 (virD2)1379309..1381414 [+], 210694310203relaxase type IV secretory pathway VirD2 
10Rmet_12601381452..1381913 [-], 46294310204hypothetical protein 
11Rmet_12611381974..1383101 [-], 112894310205putative secreted protein 
 
Region 2: 1448292..1460364
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#Locus tag (Gene)Coordinates [+/-], size (bp)Protein GIProductComponent
1Rmet_1331 (tomA0)1443460..1443684 [-], 22594310273TomA0 
2Rmet_1333 (phaB2)1444571..1445317 [+], 747943102753-oxoacyl-(acyl-carrier-protein) reductase 
3Rmet_1334 (uspA2)1445377..1445841 [+], 46594310276universal stress protein, UspA family 
4Rmet_1335 (chnA)1446018..1446806 [+], 78994310277cyclohexanol dehydrogenase 
5Rmet_1336 (traR)1446803..1447765 [+], 96394310278LysR family transcriptional regulator 
6Rmet_13371447762..1448040 [+], 27994310279lipoprotein 
7Rmet_13381447986..1448282 [-], 29794310280hypothetical protein 
8Rmet_1339 (traG)1448292..1450286 [+], 199594310281conjugal transfer coupling protein TraG  TraG
9Rmet_13401450283..1450747 [+], 46594310282CopG-like DNA-binding helix-turn-helix domain-containing protein 
10Rmet_1341 (trbB)1450744..1451808 [+], 106594310283P-type conjugative transfer ATPase TrbB  TrbB
11Rmet_1342 (trbC)1451805..1452188 [+], 38494310284conjugal transfer TrbC transmembrane protein  TrbC
12Rmet_1343 (trbD)1452185..1452457 [+], 27394310285conjugal transfer TrbD transmembrane protein  TrbD
13Rmet_1344 (trbE)1452470..1454929 [+], 246094310286conjugal transfer ATPase TrbE  TrbE
14Rmet_1345 (trbJ)1454926..1455663 [+], 73894310287conjugal transfer protein TrbJ  TrbJ
15Rmet_13461455676..1455996 [+], 32194310288lipoprotein 
16Rmet_1347 (trbL)1455993..1457372 [+], 138094310289conjugal transfer protein TrbL  TrbL
17Rmet_1348 (trbF)1457397..1458101 [+], 70594310290conjugal transfer protein TrbF  TrbF
18Rmet_1349 (trbG)1458098..1459090 [+], 99394310291conjugal transfer protein TrbG/VirB9/CagX putative exported protein  TrbG
19Rmet_1350 (trbI)1459093..1460364 [+], 127294310292type IV secretion system transmembrane protein  TrbI
20Rmet_13511460361..1460606 [+], 24694310293hypothetical protein 
21Rmet_1352 (comL)1460941..1461777 [-], 83794310294DNA uptake lipoprotein 
22Rmet_1353 (rluD)1461789..1463000 [+], 12129431029523S rRNA pseudouridine synthase 
23Rmet_13541462987..1463778 [+], 79294310296hypothetical protein 
24Rmet_13551463804..1464031 [-], 22894310297putative transmembrane protein 
 
flank Genes in the 5-Kb flanking regions if available, or non-essential genes in the T4SS gene cluster if any.

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Proteins        Genes
This T4SS is encoded by the integrative and conjugative element (ICE) CMGI-2 and is thought to contribute to the transfer of this element. (click on the ICE name to view the detail ICE informaion in ICEberg, an ICE database)
(1) Janssen PJ; Van Houdt R; Moors H; Monsieurs P; Morin N; Michaux A; Benotmane MA; Leys N; Vallaeys T; Lapidus A; Monchy S; Medigue C; Taghavi S; McCorkle S; Dunn J; van der Lelie D; Mergeay M (2010). The complete genome sequence of Cupriavidus metallidurans strain CH34, a master survivalist in harsh and anthropogenic environments. PLoS One. 5(5):e10433. [PudMed:20463976]